The shannon diversity index is a commonly used measure of diversity. Three commonly used alpha diversity measures include species observed, shannon index and simpson index. For example, species richness usually noted s of a dataset is the number of different. An example of the use of the index of diversity would be a measure of racial diversity in a city. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. But other forces greater than himself are at play, and the multilayered, complex, and nuanced on the surface, this novel tells the story of robert shannon, a former chaplain with the u. A diversity index is the measure of species diversity in a given community. H has no intuitive interpretation in terms of probability and is sensitive to sample size. H has its foundations in information theory and represents the uncertainty about the. In fact, in the late 1940s shannon had built on the work of wiener. A diversity index is a quantitative measure that reflects how many different types such as. Jan 26, 2012 a diversity index is the measure of species diversity in a given community. B10 of figure 1 of shannons diversity index except that the indices must be multiplied by 2.
It is a transmission model consisting of five elements. Species diversity is also described in terms of the phylogenetic diversity, or evolutionary relatedness, of the species present in an area. A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. How can i calculate shannons diversity index using excel sheet. Addition of menhinick, gini and strongs alpha diversity indices. Diversity index analysis tool real statistics using excel. Lets compute the shannonweiner diversity index for the same hypothetical community in the previous example. The resulting species abundance distribution, the log series model, is associated with an index of diversity known as fishers other models and measures were soon developed, and two enduring approaches are prestons log normal distribution of species abundances and. Shannonweiner index the shannon index is affected by both the number of species and their equitability, or evenness. Information theory was not just a product of the work of claude shannon. Species diversity considers both the number of species richness in the community and their relative abundance evenness or equability. Shannon s index of diversity h is derived from information theory, originally in the context of information in telephone systems shannon, 1948. Aug 28, 2014 for example, when modeling changes in species diversity throughout a community, knowledge of the genetic and chemical diversity of the primary producer e. Here is the website i show you toward the end of the.
The shannon diversity index h is another index that is commonly used to characterize species diversity in a community. Larger values reflect communities with greater species richness and evenness, while lower numbers reflect communities with fewer species andor a very uneven distribution. For example, species richness usually noted s of a dataset is the number of different species in the corresponding species list. As a measure of species diversity, we will calculate the shannon index. There have been numerous attempts to create compound indices that combine measures of richness and abundance.
Calculate the shannon diversity index and evenness for these sample values. Simpsons diversity index in a dictionary of ecology. Shannon wiener species diversity index formula data analysis. Sample values s 60,10,25,1,4 number of species n 5. Natural resource biometrics measures of diversity shannon. The shannonweiner index is most sensitive to the number of species in a sample, so it is usually considered to be biased toward measuring species richness. A diversity index is a mathematical measure of species diversity in a community. Statistics shannon wiener diversity index tutorialspoint. Pdf application of shannons index to study diversity with. It is characterized by the number of individuals observed for each species in the ecosystem. Sample values s 60,10,25,1,4 number of species n 5 first, let us calculate the sum of the given values. Find shannons index of diversity and index of relative diversity for a random sample of 25 observations distributed among five categories as shown in range b4. The simpson index is a dominance index because it gives more weight to common or dominant species.
The proportion of species i relative to the total number of species p i is calculated, and then multiplied by the natural logarithm of this proportion lnp i. Species diversity and richness 4 is designed for professional ecologists and their students. The inspiration for adopting the word entropy in information theory came from the close resemblance between shannon s formula and very similar known formulae from statistical mechanics. The same process operates with the shannon index which should anyway be replaced with exponential shannon one of the hill numbers and other diversity measures, though the problem has too. Like simpsons index, shannons index accounts for both abundance and evenness of the species present. We will then consider one measure used to compare diversity among communities, jaccard. The maximum diversity hmax of a sample is found when all species are equally abundant. Species diversity species diversity differs from species richness in that it takes into account both the numbers of species present and the dominance or evenness of species in relation to one another. The higher the value of this inverse index the greater the diversity.
In statistical thermodynamics the most general formula for the thermodynamic entropy s of a thermodynamic system is the gibbs entropy. Calculations of shannon diversity index suggest that individuals enter the sample randomly from an. A tribute to claude shannon 19162001 and a plea for. A diversity index is a quantitative measure that reflects the number of different. The shannon index increases as both the richness and the evenness of the community increase. This diversity measure is based on information theory.
However, you cannot compare the two index values using classic hypothesis tests because you do not have replicated data. Which of these diversity indices is the basic shannon index. The index is most often used for ecological studies that measure species diversity, but the same analysis can also be applied to other principles, such. What is the correct name for the biodiversity index. Diversity indices like the shannon entropy shannonwiener index and the ginisimpson index are not themselves diversities. How do i run diversity index calculation on indivi. Code to add this calci to your website just copy and paste the below code to your webpage where you want to display this calculator. As has been noted previously by some authors for example. If the community was equitable first example, wed have a 5050 chance of sampling the first species.
The first step in calculating the shannon diversity index of your closet is counting how many species life forms you have. Online biodiversity calculator which helps to calculates the shannon s diversity index and evenness from the given sample data. Shannon is sent to ireland in the summer of 1922, supposedly to trace his familys roots along the shannon river. On some occasions it is called the shannonwiener index and on other occasions it is called the shannonweaver index.
Diversity indices provide more information about community composition than simply species richness i. What does the diversity index take into account when measuring diversity. May 18, 2018 here i show how to use the shannon weiner diversity index in order to describe the variety of organisms. Simpsons diversity indexis used to calculate a measure of diversity, taking into account the number of something as well as its abundance. Here i show how to use the shannonweiner diversity index in order to describe the variety of organisms. Richness is a simple measure, so it has been a popular diversity index in ecology, where abundance data are often not available for the datasets of interest. The shannonwiener diversity index, h, is calculated using the following equation. A greater number of species and a more even distribution both increase diversity as measured by h. The information entropy, often just entropy, is a basic quantity in information theory associated to any random variable, which can be interpreted as the average level of information, surprise, or uncertainty inherent in the variables possible outcomes. If we use the compliment to simpson s d, the value is. The methods on offer range from the familiar, such as the calculation of various diversity indices and the fitting of common distributions, to more recentlydeveloped techniques such as total species complement estimators. It is often used when we wish to study a random sample from a larger community.
Shannon wiener species diversity index calculator calculate. The shannon evenness index, abbreviated as sei, provides information on area composition and richness. Shannon index the idea behind this index is that the diversity of a community is similar to the amount of information in a code or message. The shannon index has been a popular diversity index in the ecological literature. First, let us calculate the sum of the given values. The concept of information entropy was introduced by claude shannon in his 1948 paper a mathematical theory of communication. Simpson, given by d1pi2, where d is the diversity index and pi is the proportion of individuals in the ith species. In the shannon index, p is the proportion nn of individuals of one particular species found n divided by the total number of individuals found n, ln is the natural log. It is not itself diversity, but just indices of diversity. The fact that the index incorporates both components of biodiversity can be seen as both a strength and a weakness. How to calculate shannon wiener diversity index definition. Shannon index is a commonly used diversity index that takes into account.
For example, some areas may be rich in closely related taxa, having evolved from a common ancestor that was also found in that same area, whereas other areas may have an array of less closely related. Diversity indices that combine both richness and abundance i. The uncertainty is measured by the shannon function h. For our uses, this order could be characterized by the number of species andor the number of individuals in each species, within our sample plot.
Just like the diameter of a sphere is an index of its. The shannon index measures the order within a community written as h. The hutcheson ttest is a modified version of the classic ttest that provides a way to compare two samples. Introduction, simpsons index and shannonweiner index. Richness r simply quantifies how many different types the dataset of interest contains. Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. Since by definition the pis will all be between zero and one, the. It covers the number of different land cover types m observed along the straight line and their relative abundances p i. Oxford university press usa publishes scholarly works in all academic disciplines, bibles, music, childrens books, business books, dictionaries, reference books, journals, text books and more. In this second example, if we were to sample 1 individual, it likely would be of the first species.
Thus, if sunflower city was 85% white and 15% black, the index of diversity would be. Feb 12, 2016 shannonwiener diversity index in excel duration. Diversity of classes shannon, simpson, species statsdirect. The calculator uses the following formula to calculate the shannonwiener diversity index. The shannonweaver diversity index is based on communication theory.
Popular diversity books meet your next favorite book. Shannon by frank delaney meet your next favorite book. Simpson, given by d1pi2, where d is the diversity index and p i is the proportion of individuals in the ith species. The one formula that i have considered is the shannon index, but that measures the diversity with more than one species in a community, not the species attribute to the community. Note that species evenness ranges from zero to one, with zero signifying no evenness and one, a complete evenness. Here p i is the proportion of total number of species made up of the ith. They are just indices of diversity, in the same way that the diameter of a sphere is an index of its volume but is not itself the volume.
When calculating the shannon index is often used a binary logarithm, but also. Foremost among these are the shannons diversity h and simpsons diversity d 1 indices table table1, 1, which differ in their theoretical foundation and interpretation magurran 2004. The data are collected from the statistical hand book, assam, 2004 1. Shannons diversity index real statistics using excel. Divide shannons diversity index h by natural logarithm of species richness lns to calculate the species evenness. Application of shannons index to study diversity with.
Shannon in 1949 derived a function that became known as the shannon diversity index. Arguments data a list of otu tables to be processed. It was the result of crucial contributions made by many distinct individuals, from a variety of backgrounds, who took his ideas and expanded upon them. Shannons index of diversity h is derived from information theory, originally in the context of information in telephone systems shannon, 1948. A diversity index also called phylogenetic indices or phylogenetic metrics is a quantitative measure that reflects how many different types such as species there are in a dataset a community and that can simultaneously take into account the phylogenetic relations among the individuals distributed among those types, such as richness, divergence or evenness. It is calculated by dividing the shannon diversity index by its maximum h m. The document is still incomplete and does not cover all diversity methods in vegan. It combines both evenness and richness in a single measure. Indeed the diversity and directions of their perspectives and interests shaped the direction of information theory. Of the many species diversity indices used in the literature, the shannon index is perhaps most commonly used. Wiener index, shannonweaver index, species richness, species diversity. In this example, the first sample would be considered more diverse.
Like simpsons index, shannon s index accounts for both abundance and evenness of the species present. Diversity indices like the shannon entropy shannon wiener index and the ginisimpson index are not themselves diversities. Shannonwiener diversity index is a term in biologyecology. How to do the shannonweiner diversity calculation youtube.1388 1218 1220 779 379 1069 1088 1184 521 1576 1343 172 857 128 1154 325 428 884 947 1475 1233 1446 782 132 246 1318 867 1271 1254 1599 778 899 1629 1522 456 552 1640 944 996 1036 221 829 802 1321 1039 1038 214 24 555